Will PCIe x1 x4 cards work in x16 slot?

Will PCIe x1 x4 cards work in x16 slot? pcle x16 slot
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pcle x16 slot PCIe x16 Gen expansion backplane with two slots The PCIe x16 Gen3 backplane has two PCI Express x16 slots, with one slot supporting the OSS target

pcle x1 slot For example, if your graphics card is connected to a PCIe x16 slot, it means that it has 16 independent lanes dedicated just to it No other Pcie x16 slot into multiple x4 slots · PCIE Bifurcation · IBM 69Y5321 M4 PCIe Gen-III Riser Card 2 · Cablem ATIC